Scrat's continuous pursuit of an infernal acorn has world-changing consequences for Manny (Ray Romano), Sid (John Leguizamo) and Diego (Denis Leary) when the continents split apart. The continental crack-up separates Manny and his friends from the rest of the herd, including Manny's mate, Ellie (Jennifer Lopez) and rebellious daughter Peaches (Keke Palmer) . The pals are desperate to return home, but a pirate (Peter Dinklage) and his ragtag crew stand in their way.
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